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The external morphology of Pinus timleri seed cones is described from exceptionally well preserved cones and cone fragments from the Lower Rhenish Basin (Germany). The seed cones are asymmetric with a smooth side and erect umbos on the apophyses of the other side. We hypothesise that the smooth side presents the shadow-exposed side while the side with more erect umbos is the light-exposed side as in many extant Pinus seed cones.
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